Return to Skip Menu

Main Content

Thomas Fox

Honorable Garland Gray Professor of Forestry and Professor of Forest Soils and Silviculture

Ph.D., Soil Science University of Florida (1989); M.S., Forestry, Virginia Tech, 1984; Certificate of Advanced Study In Pulp and Paper Technology, University of Maine (1981); B.S., Forestry, University of Maine (1980)

Interest Areas: Forest fertilization and tree nutrition; Silviculture of Appalachian hardwoods; Intensive silviculture of southern pine plantation; Forest soils; Rhizosphere chemistry; Environmental sustainability of intensive forest management; Ecophysiology of managed forest stands.

Co-Director, Forest Productivity Cooperative
Site Director, NSF Center for Advanced Forestry Systems

Teaching Responsibilities
  • FOR 2414 Field Experiences in Forest Resources
  • FOR 3324 Silviculture Principles and Applications
  • FOR 4364 Advanced Silviculture
  • FOR 4354 Forest Soils & Hydrology
Selected Research Activities
  • Forest Productivity Research Cooperative (FPC).This is a research partnership among Virginia Tech, North Carolina State University, and the University of Concepcion in Chile that is supported by forest  industry in the United States and South America. Currently there are more than 50 forest industry organizations participating in the Cooperative which has an annual budget in excess of $800.000. The mission of the Cooperative is to integrate research, education and technology transfer to increase the productivity, profitability and sustainability of plantation forestry throughout the Americas. Dr. Fox serves as Co-Director of the Forest Productivity Cooperative.
  • NSF Center for Advanced Forestry Systems (CAFS).  The Center for Advanced Forestry Systems is a National Science Foundation Industry/University Cooperative Research Center that brings together top forestry research programs in the United States with industry members to solve complex forest management problems. The mission of CAFS is to optimize genetic and silvicultural systems to produce high-quality forest raw materials for new and existing products.  Dr. Fox serves as the Site Director for CAFS at Virginia Tech.
  • PINEMAP (Pine Integrated Network: Education Mitigation and Adaptation Project).  PINEMAP integrates research, extension, and education to enable southern pine landowners to manage forests to increase carbon sequestration; increase efficiency of nitrogen and other fertilizer inputs; and adapt forest management approaches to increase forest resilience and sustainability under variable climates. PINEMAP focuses on the 20 million acres of planted pine forests managed by private landowners in the Atlantic and Gulf coastal states from Virginia to Texas, plus Arkansas and Oklahoma. These forests provide critical economic and ecological services to U.S. citizens. Southeastern forests contain 1/3 of the contiguous U.S. forest carbon and form the backbone of an industry that supplies 16% of global industrial wood, 5.5% of the jobs, and 7.5% of the industrial economic activity of the region. Dr. Fox serves as the Silviculture and Ecophysiolgy Aim Leader and the Integration Leader for PINEMAP.
  • Use of Stable Isotopes to Evaluate Uptake and Environmental Fate of Nitrogen Fertilizers in Forest Ecosystems.   This project is using 15N labeled N fertilizers to evaluate their uptake, transformations and environmental fate in forest ecosystems.   We are comparing conventional fertilizers such as urea with enhanced efficiency fertilizers that include polymer coated urea and NBPT treated urea.  The goal of this research is to improve nutrient use efficiency and reduced loses through volatilization and leaching following fertilization of forest ecosystems. Research installations are located throughout the South in loblolly pine plantations.
  • Application of Biosolids to Forests.We are working with the Washington DC Council of Governments to evaluate the impacts of biosolids applications on forest growth, nutrient dynamics and water quality in Virginia.
  • Restoring Sustainable Forests on Appalachian Strip Mined Lands.We are working to restore productive forests to lands that were strip mined for coal in Virginia, West Virginia and Ohio. Prior to mining, most of these lands supported stands of mixed hardwoods. We are comparing site preparation, planting and subsequent management practices required to successfully reestablish white pine, hybrid poplar and mixed stands of native hardwoods on reclaimed strip mines. This work is in cooperation with the Powell River Project.
  • Impact of Weed Control and Fertilization on Growth of Pine Plantations.This project is evaluating the effect of intensive management on the growth of White pine, shortleaf pine, Virginia pine and Loblolly pine plantains in the Virginia Piedmont. A factorial combination of weed control and annual fertilization is being applied to these stands. This study is being conducted at the Virginia Tech Reynolds Homestead Research Center.
  • The Effects of Alternative Regeneration Methods on Hardwood Regeneration in Natural Hardwood Stands the Appalachian Region. We are evaluating the effects of alternative silvicultural systems including clearcuting, shelterwood, leave-tree, and group selection on species composition and growth of upland hardwoods in the Appalachian Mountains of Virginia and West Virginia. The goal of this project is to develop sustainable forest management practices for the Appalachian hardwood forests. It is a cooperative project with USDA Forest Service, Southern Research Station and the Bent Creek Experiment Forest.
Professional Achievements
  • 18 years of industrial forestry experience in New England, the Pacific Northwest and The South
  • Registered Professional Forester in Georgia and Maine
  • Certified Professional Soil Scientist
  • Recipient of Florida SAF Stephen Spurr Award for Forestry Research
  • Associate Editor for Silviculture of Southern Journal of Applied Forestry (1993-1998)
  • Chair of Southern Industrial Forest Research Council (1999-2000)
  • Co-Chair of National Council for Air and Stream Improvement Forest Sustainability Task Group (1995-2000)
  • Chair SAF Forest Soils Working Group
  • Chair Suwannee Chapter of Florida SAF
  • Forest Science and Technology Chair for Appalachian SAF
  • Authored or co-author over 75 technical forestry publications
  • SAF Certified Forester
  • Associate Editor - Soil Science Society of America Journal (2000 - present)
Referred Publications

Book Chapters

  • Comerford, N.B. and T.R. Fox. 2015. Forest Soils.  In Encyclopedia of Soil Science. Marcel Dekker, Inc. New York.
  • Fox, T.R. and J. Creighton. 2015. Silvics of Oak.  In A Practical Guide to Managing Oak Forests in the Eastern United States. CRC  Press. Boca Raton, FL. 
  • Rubilar, R.A., H.L. Allen, T.R. Fox, J.L. Stape, and T.J. Albaugh. 2015. Disponibilidad de nutrients y fertilizacion en plantaciones forestales. In: Fertilidad de suelos y fertilizacion de cultivos. Editors: Echeverria, H.E. and Garcia, F.O. INTA Ediciones and IPNI Cono Sur. Buenos Aires, Argentina. Pages 753-769.
  • Kiser, L.C. and T.R.  Fox. 2013.  Short Rotation Woody Crop Biomass Production for Energy.  Chapter 6. pp. 205-237.  In B. Singh (Ed.). Biofuel Crop Sustainability. Blackwell Science Publishers. Oxford, England.
  • Fox, T.R., Miller, BH, Stape, J.L., Rubilar, R.P. and Albaugh, T.J. 2010. Phosphorus Nutrition and Fertilization in Forest Plantations.  In Bunemann, E., Oberson, A. and Frossard, E. (eds.) Phosphorus in Action – Biological Processes in Soil Phosphorus Cycling.  Soil Biology Series. Springer.
  • Fox, T.R., and R. R. Hicks. 2004. Forest Productivity. Chapter 6. In Southern Forest Science: Past, Present, Future. USDA Forest Service. General Technical Report SRS-75. 394 pp.
  • Fox, T.R., E. Jokela, and H.L. Allen. 2004. The evolution of pine plantations in the southern United States. Chapter 8. In Southern Forest Science: Past, Present, Future. USDA Forest Service. General Technical Report SRS-75. 394 pp.
  • Fox, T.R. 2001. Sustained productivity in intensively managed forest plantations. pp. 187-202 In Forest Soils and Ecosystem Sustainability. Elsevier. Amsterdam. 462 pp.
  • Fox, T.R. 1995. Low molecular weight organic acids in forest soils: Influence on metal solubility and nutrient availability. pp. 43-62 In Carbon Forms and Functions in Forest Soils. SSSA. Madison, WI.
  • Graham, R.L., T.R. Fox, and P.H. Dougherty. 1986. Multiple stress factors: The potential role of systems models in assessing the impact of multiple stresses on forest productivity. Chapter 8. In Stress Physiology and Forest Productivity. Martinus Nijhoff Pub., Dordrecht, Netherlands. 239 pp.

Papers in Refereed Journals

  • Subedi, S., and T.R. Fox. 2016. Predicting Loblolly Pine Site Index from Soil Properties Using Partial Least Square Regression. Forest Science
  • Sumnall, M.J., T.R. Fox, R.H. Wynne, V.A. Thomas. 2016. Analysis of a LiDAR voxel-derived vertical profile at the plot and the individual tree scale for the estimation of forest canopy layer characteristics.  International Journal of Remote Sensing. In Press.
  • Sumnall, M.J., A. Peduzzi, T.R. Fox, R.H. Wynne, V.A. Thomas, B. Cook. 2016. Assessing the transferability of statistical predictive models for leaf area index between two airborne discrete return LiDAR sensor designs within multiple intensively managed Lobolly pine forest locations in the south-eastern USA. Remote Sensing of Environment. 176:308-319.
  • Ramirez, M.V., R.A. Rubilar, C. Montes, J.L. Stape, T.R. Fox, and H. L. Allen. 2016. Nitrogen availability and mineralization in Pinus radiata stands fertilized midrotation at three contrasting sites. Journal of Soil Science and Plant Nutrition. doi: 10.4067/S0718-95162016005000009.
  • Minnick, K.J., C.B. Pandey, and T.R. Fox. 2016. Dissimilatory nitrate reduction to ammonia and denitrification: Effect of soil moisture and N fertilization regimes in loblolly pine forests. Biology and Fertility of Soils. In Press. 
  • Sumnall, M.J., T.R.Fox, R.H. Wynne, C. Blinn, and V.A. Thomas. 2015. Estimating leaf area index at multiple heights within the understory component of Loblolly pine forests from airborne discrete-return lidar. International Journal of Remote Sensing. 37(1):78-99. Doi. 10.1080/01431161.2015.1117683.
  • Subedi,S., T.R. Fox, and R.H. Wynne. 2015. Determination of Fertility Rating (FR) in the 3-PG Model for Loblolly Pine Plantations in the Southeastern United States Based on Site Index. Forests.  6:3002-3027; doi:10.3390/f6093002
  • Albaugh, T. J., T.R. Fox, H.L. Allen, and R.A. Rubilar. 2015. Juvenile Southern Pine Response to Fertilization Is Influenced by Soil Drainage and Texture. Forests. 6: 2799-2819; doi:10.3390/f6082799
  • Yanez, M.A., T.R. Fox, and J.R. Seiler. 2015. Early growth response of loblolly pine varieties and families to silvicultural intensity. Forest Ecology and Management. 356:204-215.
  • Will, R., T.R. Fox, M. Akers, J.C. Domec, C. Gonzalez-Benecke, E. J. Jokela,  M. Kane,M.A. Laviner, G. Lokuta, D. Markewitz, M.A. McGuire, C.Meek, A. Noormets, L. Samuelson,  J. Seiler, B. Strahm, R. Tesky, J. Vogel, E. Ward, J. West, D. Wilson, T. A. Martin. 2015. A Range-wide Experiment to Investigate Nutrient and Soil Moisture Interactions in Loblolly Pine Plantations. Forests. 2015, 6(6), 2014-2028; doi:10.3390/f6062014
  • Albaugh, T.J., J. Alvarez, R. Rubilar, T.R. Fox, H.L. Allen, J. L. Stape, and O. Mardones. 2015. Long-term Pinus radiata productivity gains from tillage, vegetation control and fertilization. Forest Science. 61(4):800–808.
  • Albaugh, T.J., R.A. Rubilar, T.R. Fox, H.L. Allen, J.B. Urrego, M Zapata, and J. L. Stape. 2015. Response of Eucalyptus grandis in Colombia to mid-rotation fertilization is dependent on site and rate but not frequency of application. Forest Ecology and Management. 350:30-39.
  • Ward, E.J., J.C.,Domec, M.A.  Laviner,  T.R. Fox, Ge Sun, S.  McNulty, J.  King, and A. Noormets, 2015. Fertilization intensifies drought stress: Water use and stomatal conductance of Pinus taeda in a midrotation fertilization and throughfall reduction experiment. Forest Ecology and Management.355:72-82
  • Minick, K.J., Strahm, B.D., Fox, T.R., Sucre, E. B., and Leggett, Z. H. 2015.  Microbial Nitrogen Cycling Response to Forest-Based Bioenergy Production in a Loblolly Pine Forest in Coastal North Carolina. Ecological Applications. 25(8):2366-2381.
  • Rubilar, R., T.Fox, J. Stape, T. Albaugh. 2014. Limitantes para la productividad potencial de pinus y eucalyptus en Sudamerica. XXVIII Jornadas Forestales de Entre Rios.  Concordia, 2014:1-10.
  • Gonzalez-Benecke, C., Gezan, S.,  Albaugh, T. Allen, H. Burkhart, H. Fox, T. Jokela, E., Meier, C. Martin, T., Rubilar, R., and Saumelson, L. 2014. Local and general above-stump biomass functions for  loblolly pine and slash pine trees. Forest Ecology and Management. 334:254-276.
  • Elliot, J.R. and T.R. Fox. 2014. Ammonia volatilization following fertilization with urea or ureaform in a thinned loblolly pine plantation. Soil Science Society of America Journal.
  • Carlson, C.A., T.R. Fox, H. L. Allen, T.J. Albaugh, J.L. Stape, and R.A. Rubilar. 2014. Impacts of combination plowing on the survival and growth of loblolly pine. Tree Planters’ Notes. 57 (7):18-22.
  • Raymond, J., T.Fox, and B.Strahm. 2014. Can enhanced efficiency fertilizers affect the fate of nitrogen in loblolly pine plantations.  Better Crops. 98 (2014, No. 2) 4-6.
  • Minnick, K.J., B.D. Strahm, T.R. Fox, E.B. Sucre, Z.H. Leggett, J.L. Zerpa. 2014. Switchgrass intercropping reduces soil inorganic nitrogen in a young loblolly pine planation located in coastal North Carolina. Forest Ecology and Management. 319 (2014):161-168.
  • Albaugh, T.J., L.C. Kiser, T.R. Fox, H.L. Allen, R.A. Rafael, and J. L. Stape. 2014. Ecosystem nutrient retention after fertilization of Pinus taeda. Forest Science. 10.5489/forsci.13-159.
  • Watt, M., R. Rubilar, M. Kimberley, D. Kriticos, V. Emhart, O. Mardones, M.Acevedo, M. Pincheira, J. Stape, T. Fox. 2014. Using seasonal measurements to inform ecophysiology; Extracting cardinal growth temperatures for process based growth models of five Eucalyptus  species/crosses  from simple field trials.  New Zealand Journal of Forestry Science. 44:9.
  • Carlson, C.A., T.R. Fox, H.L. Allen, T.A. Albaugh, J.L. Stape, and R.P. Rubilar. 2014. Growth responses of loblolly pine in the Southeast United States to midrotation applications of nitrogen, phosphorus, potassium and micronutrients. Forest Science. 60(1): 157-169.
  • Rubilar, R.A., T. J. Albaugh, H.L. Allen, J. Alvarez, T.R. Fox, and J. L. Stape. 2013.  Foliage development and leaf area duration in Pinus radiata.  Forest Ecology and Management.  304:455-463.
  • Stanturf, J.A., E.D. Vance, T.R. Fox, and M. Kirst. 2013. Eucalyptus beyond its native range: Environmental issues in exotic bioenergy plantations. International Journal of Forestry Research. Doi: 10.1155/2013/463030.
  • Kiser, L.C. T.R. Fox, and C.A. Carlson. 2013. Foliage and litter chemistry, decomposition, and nutrient release in Pinus taeda. Forests. 4:137-154. Doi:10.3390/f4010137
  • Albaugh, T.J., T.R. Fox, C. A. Blinn,  H.L. Allen, R. Rubiliar, J.L. Stape, 2013. Developing a new foliar-nutrient based method to predict response to competing vegetation control in Pinus taeda. Southern Journal of Applied Forestry. 37(4): 196-200.
  • Sabatia, C.A., T.R. Fox and H. Burkhart. 2013. Extending a model system to predict biomass in mixed-species southern Appalachian hardwood forests. Southern Journal of Applied Forestry. DOI 10.5849/sjaf.12-005. 37(2): 122-126.
  • Campoe, O.C., J.L. Stape, T. J. Albaugh, H. L. Allen, T.R. Fox, R. Rubilar, and D. Binkley. 2013.  Fertilization and irrigation effects on tree level aboveground net primary production, light interception and light use efficiency in a loblolly pine plantation. Forest Ecology and Management. 288:43-48.
  • Rubilar, R., T.J. Albaugh, H.L. Allen, J. Alvarez, T.R. Fox, and J.L. Stape. 2013. Influences of silvicultural manipulations on above- and below-ground biomass accumulation and leaf area in young Pinus radiata plantations at three contrasting sites in Chile.  Forestry.86:27-38.
  • Stovall, J.P., J.R. Seiler, and T.R. Fox. 2013.  Respiratory C fluxes and root exudation differ in two full-sib clones of Pinus taeda (L.) under contrasting fertilizer regimes.  Plant and Soil. 363:257-271.
  • Stovall, J.P., J.R. Seiler, and T.R. Fox. 2013. Allometry varies among six-year-old Pinus taeda (L.) clones in the Virginia Piedmont. Forest Science. 59(1):60-62.
  • Jones, P.D. and T.R. Fox. 2012. Stem sinuosity in young and merchantable stands of Pinus taeda Forest Products Journal.62:354-358.
  • Albaugh, T.J., E.D. Vance, C. Gaudreult, T.R. Fox, H. L. Allen, J. L. Stape, and R.A. Rubilar.  2012. Carbon emissions and sequestration from fertilization of pine in the southeastern United States. Forest Science. 58:419-429.
  • Albaugh, T.J., H.L. Allen, J.L. Stape, T.R.Fox, R.A. Rubilar, and J. Price. 2012. Intra-annual nutrient flux in Pinus taeda. Tree Physiology. 32:1237-1258
  • Kiser, L.C. and T.R. Fox. 2012. Nitrogen and phosphorus pools in fertilized loblolly pine and sweetgum. Soil Science Society of America Journal.76:2278-2288.
  • Stovall, J.P., T.R. Fox, and J. R. Seiler. 2012.  Short-term changes in biomass partitioning of two full-sib clones of Pinus taeda L. under differing fertilizer regimes over four months. Trees: Structure and Function. 26:951-961.
  • Peduzzi, A., R.H. Wynne, T.R. Fox, R.F. Nelson and V.A. Thomas. 2012. Estimating leaf area index in intensively managed pine plantations using airborne laser scanner data. Forest Ecology and Management. 270:54-65.
  • Blinn, C.E., T.J. Albaugh, T.R. Fox, R.H. Wynne, J.L. Stape, R.A. Rubilar and H.L. Allen. 2012. A Method for Estimating Deciduous Competition in Pine Stands using Landsat. Southern Journal of Applied Forestry. 36:71-78.
  • Vickers, L.A., T.R. Fox, D. L. Loftis, and D. A. Boucugnani. 2012.  Predicting Forest Regeneration in the Central Appalachians Using the REGEN Expert System. Journal of Sustainable Forestry.30 (8):790-822.
  • Atwood, C.J., T.R. Fox, and D. L. Loftis. 2011.  Effects of Alternative Silvicultural Treatments on Regeneration in the Southern Appalachians. Journal of Sustainable Forestry.  30:419-440.
  • Harrison, R.H, D. Richter, and T. R. Fox. 2011.  Deep soils.  Forest Science.  57:1-2.
  • Sucre, E.B. J.W. Tuttle, and T.R. Fox. 2011. The use of ground-penetrating radar as a tool to accurately estimate soil depth in rocky forest soils of the southern Appalachians. Forest Science. 57:59-66.
  • Miller, B.W. and T.R. Fox. 2011. Long-term Fertilizer Effects on Oxalate Desorbable Phosphorus Pools in a Typic Paleaquult. Soil Science Society of America Journal. 75 (3):1110-1116.
  • Zerpa, J. L. and T.R. Fox. 2011. Controls of Volatile NH3 losses from loblolly pine plantations fertilized with urea in the Southeast US. Soil Science Society of America Journal. 75:257-266.
  • Stovall, J. L., C.A. Carlson, J.R. Seiler, T.R. Fox, and M.A. Yanez. 2011. Growth and stem quality responses to fertilizer applications by 21 loblolly pine clones in the Virginia Piedmont. Forest Ecology and Management. 261: 362-372.
  • Rubilar, R.A., H.L. Allen, J.S. Alvares, T.J. Albaugh, T.R. Fox, and J.L. Stape 2010. Silvicultural manipulation and site effect on above and below ground biomass equations for young Pinus radiata plantations. Biomass and Bioenergy 34:1825-1837.
  • Munsell, J.F., and T.R. Fox. 2010. An analysis of the feasibility for increasing woody biomass production from pine plantations in the southern United States. Biomass and Bioenergy.34:1631-1642.
  • Homyack, J.A., E.B. Sucre, C.A. Haas, and T.R. Fox. 2010. Does Plenthodon cinereus affect leaf litter decomposition and invertebrate abundances in mixed oak forests? Journal of Herpetology.44:447-456.
  • Albaugh, T.J., H.L. Allen, J. Stape, T.R. Fox, R.A. Rubilar, C.A. Carlson, and R. Pezutti. 2010. Leaf area duration in natural range and exotic Pinus taeda. Canadian Journal of Forest Science. 40:224-234.
  • Albaugh, J., H.L. Allen, T.J. Albaugh, T.R. Fox, J.L. Stape, and R.P. Rubilar. 2010. Characterization of foliar macro- and micro-nutrient concentrations and ratios in loblolly pine plantations in the southeastern United States. Southern Journal of Applied Forestry. 34(2):53-64.
  • Pratt, W.A. and T.R. Fox. 2009. Streamside management zone effectiveness for protecting water quality following forestland application of biosolids. Journal of Environmental Quality.  38:2106-2120.
  • Sucre, E.B. and T.R. Fox. 2009. Contribution of Soil Influenced by Decomposing Stumps and Root Systems to Soil Nutrient Pools in Southern Appalachian Hardwood Forests. Forest Ecology and Management. 258: 2242-2248
  • Carlson, C.A., T.R. Fox, H.E. Burkhart, H.L. Allen, and T. J. Albaugh. 2009. Accuracy of subsampling for height measurements in loblolly pine plots. Southern Journal of Applied Forestry. 33(3):145-149.
  • Albaugh, T.J., H.L. Allen, T.R. Fox, C.A. Carlson, and R. Rubilar. 2009. Opportunities for fertilization of loblolly pine in the sandhills of the southeastern United States. Southern Journal of Applied Forestry. 33(3):129-136.
  • Carlson, C.A., T.R. Fox, J. Creighton, P. M. Dougherty, and J.R. Johnson. 2009. Nine year growth responses to planting density manipulation and repeated early fertilization in a loblolly pine plantation in the Virginia Piedmont. Southern Journal of Applied Forestry. 33(3):109-114.
  • Atwood, C.J., T.R. Fox, and D.L. Loftis. 2009. Effects of alternative silviculture on stump sprouting in the Southern Appalachians. Forest Ecology and Management. 257:1305-1313.
  • Carlson, C.A., T.R. Fox, H. L. Allen, and T.J. Albaugh. 2008. Modeling mid-rotation fertilizer responses using the age-shift approach. Forest Ecology and Management.
  • Carlson, C.A., H. E. Burkhart, T. R. Fox, and H. L. Allen. 2008. Changes to the diameter distribution of Pinus taeda as a result of midrotation fertilizer applications. Canadian Journal of Forest Research. 38:2063-2071.
  • Albaugh, T.A., H.L. Allen and T.R. Fox. 2008. Nutrient use and uptake in Pinus taeda. Tree Physiology. 28:1083-1098.
  • King, N.T., J. R. Seiler, T. R. Fox, and K. H. Johnsen.  2008.  Post-fertilization loblolly pine clone physiology and growth performance. Tree Physiology 28:703-711.
  • Tyree, M.C., J.R. Seiler, and T.R. Fox. 2008. The effects of fertilization on soil respiration in two-year-old Pinus taeda L. clones. Forest Science, 54(1): 21-30.
  • Fox, T.R., H. L. Allen, T.J. Albaugh, C.A. Carlson, R. Rubilar.2007.  Forest fertilization and water quality in the United States. Better Crops. 91(2007-1): 5-7.
  • Fox, T.R., E.J. Jokela, and H.L. Allen. 2007. The development of pine plantation silviculture in the southern United States. Journal of Forestry. 105(5):337-347.
  • Jones, P.D., and T.R. Fox. 2007. Wood density in Pinus taeda x Pinus rigida and response 10 years after thinning in Virginia. Forest Products Journal. 57(12):70-73.
  • Fox, T.R., H.L. Allen, T.J. Albaugh, R. Rubilar, and C.A. Carlson. 2007. Tree nutrition and forest fertilization of pine plantations in the southern United States. Southern Journal of Applied Forestry. 31(1): 5-11.
  • Albaugh, T.A., H.L. Allen, and T.R. Fox. 2007. Historical patterns of forest fertilization in the southern United States from 1969 to 2004. Southern Journal of Applied Forestry. 31(3):129-137.
  • Tyree, M.C., J.R. Seiler, W.M. Aust, D.A. Sampson, and T.R. Fox. 2006. Long-term effects of site preparation and fertilization on total soil CO2 efflux and heterotrophic respiration in a 33-year-old Pinus taeda L. plantation on the wet flats of the Virginia Lower Coastal Plain.  Forest Ecology and Management. 234(1-3):363-369.
  • Casselman, C.N. T.R. Fox, and J.A. Burger. 2006. Thinning response of a white pine stand on a reclaimed surface mine in southwestern Virginia. Northern Journal of Applied Forestry. 24(1):9-14.
  • Carlson, C.A., T.R. Fox, S.R. Colbert, D.L. Kelting, H. L. Allen, and T.J. Albaugh. 2006. Growth and survival of Pinus taeda in response to surface and subsurface tillage in the southeastern United States. Forest Ecology and Management. 234(1-3):209-217.
  • Albaugh, T.J., H.L. Allen, and T.R. Fox. 2006. Individual tree crown and stand development in Pinus taeda under different fertilization and irrigation regimes.  Forest Ecology and Management: 234(1-3):10-23.
  • Eisenbies, M.H., J.A. Burger, W.M. Aust, S.C. Patterson, and T.R. Fox. 2006. Assessing change in soil site productivity of intensively managed loblolly pine plantations. Soil Science Society of America Journal. 70: 130-140.
  • Casselman, C.N., T.R. Fox, J.A. Burger, A.T. Jones, and J. M. Galbraith. 2006. Effects of silvicultural treatments on survival and growth of trees planted on reclaimed mine lands in the Appalachians. Forest Ecology and Management. 223(1-3):403-414.
  • Amishev, D.Y. and T.R. Fox. 2006. The effect of weed control and fertilization on survival and growth of four pine species in the Virginia Piedmont.  Forest Ecology and Management. 236(1):93-101.
  • Kyle, K.H., L.J. Andrews, T.R. Fox, W.M. Aust, J.A. Burger and G. H. Hansen. 2005.  Long-term impact of drainage, bedding, and fertilization on growth of loblolly pine (Pinus taeda L.) in the Coastal Plain of Virginia.  Southern Journal of Applied Forestry. 29(4):205-214.
  • Fisher, R.F., T.R. Fox, T. Terry and R. Harrison. 2005. Forest soils education and research: Trends, needs and wild ideas. Forest Ecology and Management. 220(1-3):1-16.
  • Allen, H.L., T.R. Fox and R.G. Campbell. 2005. What’s ahead for intensive pine plantation silviculture? Southern Journal of Applied Forestry. 29:62-69.
  • Fox, T.R. 2004. Nitrogen mineralization following nitrogen fertilization of Douglas fir forests in western Washington. Soil Science Society of America Journal. 68:1720-1728.)
  • Fox, T.R., and R.E. Kreh. 2003. Growth response of pitch x loblolly hybrid pine following crown touching release. Northern Journal of Applied Forestry. 20(4):161-166.
  • Johnsen, K., L. Samuelson, R. Teskey, S. McNulty, and T. R. Fox. 2001. Process models as tools in forestry research and management. Forest Science. 47:2-8.
  • Fox, T.R. 2000. Sustained productivity of intensively managed forest plantations. Forest Ecology and Management. 138(1-3):187-202.
  • Lan, M., N.B. Comerford, and T.R. Fox. 1995. Effects of organic anions with different Al complexation constants on P release from Spodic horizons. Soil Science Society of America Journal. 59:1745-1749.
  • Fox, T.R., and N.B. Comerford. 1992. Influence of oxalate loading on phosphorus and aluminum solubility in Spodosols. Soil Science Society of America Journal. 56:290-294.
  • Fox, T.R., and N.B. Comerford. 1992. Rhizosphere phosphatase activity and phosphatase hydrolyzable organic phosphorus in two forested Spodosols. Soil Biology and Biochemistry. 24:579-583.
  • Maimone, R.A., L.A. Morris, and T.R. Fox. 1991. Soil nitrogen mineralization potential in a fertilized loblolly pine plantation. Soil Science Society of America Journal. 55:522-527.
  • Fox, T.R., N.B. Comerford, and W.W. McFee. 1990. Phosphorus and aluminum release from a spodic horizon mediated by organic acids. Soil Science Society of America Journal. 54:1763-1767.
  • Fox, T.R., N.B. Comerford, and W.W. McFee. 1990. Kinetics of Phosphorus release from Spodosols: Effects of oxalate and formate. Soil Science Society of America Journal. 54:1441-1447.
  • Fox, T.R., and N.B. Comerford. 1990. Low-molecular-weight organic acids in selected forest soils of the southeastern USA. Soil Science Society of America Journal. 54:1139-1144.
  • Fox, T.R. 1986. Raspberry (Rubus ideaus L.) competition effects on balsam fir (Abies balsamea (L.) Mill.) seedlings in northern Maine. Tree Planters' Notes. 1986:20-23.
  • Fox, T.R., J.A. Burger, and R.E. Kreh. 1986. Effects of site preparation on nitrogen dynamics in the southern Piedmont. Forest Ecology & Management. 15(4):241-256.
Professional Accomplishments and Awards


  • ARCPACS Certified Professional Soil Scientist (Certification No. 02114)
  • SAF Certified Forester (Certification No. 1637)
  • Licensed Professional Forester - Maine  (License No. 650)
  • Registered Professional Forester - Georgia  (Registration No. 2182)


  • IUFRO Forest Soils Working Group Chair
  • IUFRO Planted Forest Task Force
  • Society of American Foresters Member since 1976
  • Society of American Foresters Silviculture Instructors Working Group
  • Society of American Foresters APSAF Science and Technology Chair
  • Society of American Foresters Soils Working Group Chair
  • Society of American Foresters Florida Suwannee Chapter Chair
  • Soil Science Society of America Member Since 1982
  • Soil Science Society of America Board of Directors
  • Soil Science Society of America Forest Soils Division Chair
  • Soil Science Society of America Reorganization Task Force Chair
  • Virginia Department of Forestry Research Advisory Committee
  • Virginia Forestry Association, Board of Directors
  • Georgia Traditional Industries Program - Fiber Supply Working Group Chair
  • Georgia Forestry Association Environmental Committee
  • Florida Forestry Association Endangered Species Task Force
  • Florida Forestry Association Forest Fertilization & Water Quality Task Force
  • Associate Editor – Forest Science
  • Associate Editor – Soil Science Society of America Journal
  • Associate Editor – Southern Journal of Applied Forestry
  • Guest Associate Editor – Forest Science
  • Guest Associate Editor – International Journal of Forestry
  • Guest Associate Editor – New Forests
  • Editorial Board – Revista BOSQUE
  • AF&PA Sustainable Forestry and Wetlands Committees
  • AF&PA Forest Science and Technology Committee
  • NCASI Forest Environmental Studies Task Group (Chair)
  • NCASI Forest Health and Wetlands Research Committee (Chair)
  • Florida Division of Forestry BMP Technical Advisory Committee        


  • University of Maine SFR Distinguished Alumni Award (2016)
  • SAF Barrington Moore Award (2013)
  • Fellow Soil Science Society of America (2012)
  • Fellow Society of American Foresters (2012)           
  • Southern Extension Forest Resources Specialists Gold Award (2012)
  • Virginia Forestry Association Certificate of Appreciation (2011)
  • Fulbright Scholar (2010-2011)
  • Virginia Tech Scholar of the Week (2007, 2014)
  • Virginia Tech College of Natural Resources Award for Outreach Excellence (2006)
  • H.J. Heinz III Center for Science, Economics and the Environment Certificate of Appreciation (2004)
  • Soil Science Society of America Service Award (2004)
  • NCASI Outstanding Service Award (2000)
  • Steven Spurr Research Award Florida Society of American Foresters (1998)
  • Rayonier President’s Achievement Award (1998)
  • University of Florida Forest Biology Research Cooperative Recognition Award (1997)
  • Who's Who in Science and Engineering (1990 to present)
  • Sigma Xi (1989); Xi Sigma P (1980);  Alpha Zeta (1979)     

About Us

Contact Us

About Cheatham Hall 


FREC Strategic Plan


2015 FREC Annual Report


Fall 2016 FREC Newsletter

View Archive Newsletters 


FREC Career Book


FREC Course Cards

FREC YouTube Videos

FREC Undergraduate Checksheets

Undergraduate Brochure (pdf)

Graduate Brochure (pdf)